❓ 题目 According to the passage, what was Robert Paine's main contribution to ecology?
- A. He discovered that physical factors are the primary determinants of ecosystem structure.
- B. He provided experimental evidence that certain species can have a disproportionate impact on biodiversity.
- C. He was the first to propose that all species are equally important in maintaining ecosystem function.
- D. He demonstrated that removing a predator always increases species diversity in intertidal zones.